If the food box is empty and a citizen still must eat, a citizen starves and the population count will be reduced by one. If the city is not producing enough food to meet demand, this food will be taken from the food box. A citizen will never take more or less than this. Each citizen requires two food per turn to survive. The food box grows with the size of the city, (and thus requires ever more surplus food to fill it in order to get the next citizen or worker), usually at a constant rate. When it is filled, another citizen will be added to the city and the food box will empty. Surplus food will accumulate until the food box is full.
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